حوت أحدب
Humpback whale [1] | |
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الحجم مقارناً بإنسان متوسط | |
التصنيف العلمي | |
مملكة: | |
Phylum: | |
Class: | |
Subclass: | |
Order: | |
Suborder: | |
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Genus: | Megaptera گرِيْ، 1846
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Species: | M. novaeangliae
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Binomial name | |
Megaptera novaeangliae بوروڤسكي، 1781
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Subspecies | |
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نطاق الحوت الأحدب (بالأزرق) | |
Synonyms | |
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' الحوت الأحدب (Megaptera novaeangliae) هو نوع من الحوت بالين. وهو rorqual (أحد أعضاء فصيلة Balaenopteridae) وهو النوع الوحيد في جنس Megaptera. ويتراوح طول البالغين من 14-17 متر والوزن يصل إلى 40 طن متري. الأحدب له شكل جسم مميز، مع زعانف صدرية طويلة و حديبات على رأسه. ويـُعرف باختراقه سطح الماء وألعابه المميزة الأخرى على سطح الماء، مما يجعله ذا شعبية مع مشاهدي الحيتان. تستطيع الذكور انتاج غناءً معقداً يستمر، عادةً، من 4 إلى 33 دقيقة.
Found in oceans and seas around the world, humpback whales typically migrate up to 16,000 km (9,900 mi) each year. They feed in polar waters and migrate to tropical or subtropical waters to breed and give birth. Their diet consists mostly of krill and small fish, and they use bubbles to catch prey. They are promiscuous breeders, with both sexes having multiple partners. Orcas are the main natural predators of humpback whales.
Like other large whales, the humpback was a target for the whaling industry. Humans once hunted the species to the brink of extinction; its population fell to around 5,000 by the 1960s. Numbers have partially recovered to some 135,000 animals worldwide, while entanglement in fishing gear, collisions with ships, and noise pollution continue to affect the species.
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التصنيف
The humpback was first identified as baleine de la Nouvelle Angleterre by Mathurin Jacques Brisson in his Regnum Animale of 1756. In 1781, Georg Heinrich Borowski described the species, converting Brisson's name to its Latin equivalent, Balaena novaeangliae. In 1804, Bernard Germain de Lacépède shifted the humpback from the family Balaenidae, renaming it B. jubartes. In 1846, John Edward Gray created the genus Megaptera, classifying the humpback as Megaptera longipinna, but in 1932, Remington Kellogg reverted the species names to use Borowski's novaeangliae.[3] The common name is derived from the curving of their backs when diving. The generic name Megaptera from the Ancient Greek mega- μεγα ("giant") and ptera/ πτερα ("wing")[4] refer to their large front flippers. The specific name means "New Englander" and was probably given by Brisson due to regular sightings of humpbacks off the coast of New England.[3]
Humpback whales are rorquals, members of the Balaenopteridae family, which includes the blue, fin, Bryde's, sei and minke whales. A 2018 genomic analysis estimates that rorquals diverged from other baleen whales in the late Miocene, between 10.5 and 7.5 million years ago. The humpback and fin whale were found to be sister taxon (see phylogenetic tree below).[5] There is reference to a humpback-blue whale hybrid in the South Pacific, attributed to marine biologist Michael Poole.[6][7]
Balaenopteridae |
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Modern humpback whale populations originated in the southern hemisphere around 880,000 years ago and colonized the northern hemisphere 200,000–50,000 years ago. A 2014 genetic study suggested that the separate populations in the North Atlantic, North Pacific, and Southern Oceans have had limited gene flow and are distinct enough to be subspecies, with the scientific names of M. n. novaeangliae, M. n. kuzira and M. n. australis respectively.[8] A non-migratory population in the Arabian sea has been isolated for 70,000 years.[9]
الوصف
The adult humpback whale is generally 14–15 m (46–49 ft), though longer lengths of 16–17 m (52–56 ft) have been recorded. Females are usually 1–1.5 m (3 ft 3 in – 4 ft 11 in) longer than males.[10] The species can reach body masses of 40 metric tons (44 short tons). Calves are born at around 4.3 m (14 ft) long with a weight of 680 kg (1,500 lb).[11]
The body is bulky with a thin rostrum and proportionally long flippers, each around one-third of its body length.[12][13] It has a short dorsal fin that varies from nearly non-existent to somewhat long and curved. As a rorqual, the humpback has grooves between the tip of the lower jaw and the navel.[10] They are relatively few in number in this species, ranging from 14–35.[12] The mouth is lined with baleen plates, which number 270-400 for both sides.[13]
Unique among large whales, humpbacks have bumps or tubercles on the head and front edge of the flippers; the tail fluke has a jagged trailing edge.[10][13] The tubercles on the head are 5–10 cm (2.0–3.9 in) thick at the base and poke up to 6.5 cm (2.6 in). They are mostly hollow in the center, often containing at least one fragile hair that erupts 1–3 cm (0.39–1.18 in) from the skin and is 0.1 mm (0.0039 in) thick. The tubercles develop early in the womb and may have a sensory function as they are rich in nerves.[14]
The dorsal or upper-side of the animal is generally black; the ventral or underside has various levels of black and white coloration.[10] Whales in the southern hemisphere tend to have more white pigmentation. The flippers can vary from all-white to white only on the undersurface.[11] The varying color patterns and scars on the tail flukes distinguish individual animals.[15][16] The end of the genital slit of the female is marked by a round lobe. This lobe visually distinguishes males and females.[13]
السلوك والبيئة
Humpback whale groups, aside from mothers and calves, typically last for days or weeks at the most.[10][17] They are normally sighted in small groups though large aggregations form during feeding and among males competing for females.[17] Humpbacks may interact with other cetacean species, such as right whales, fin whales, and bottlenose dolphins.[18][19][20] Humpbacks are highly active at the surface, performing aerial behaviors such as breaching and surface slapping with the tail (lobtailing) and flippers. These may be forms of play and communication and/or for removing parasites.[10]
Humpbacks rest at the surface with their bodies laying horizontally.[21] The species is a slower swimmer than other rorquals, cruising at 7.9–15.1 km/h (4.9–9.4 mph). When threatened, a humpback may speed up to 27 km/h (17 mph).[13] They frequent shallow seamounts, commonly exploring depths of up to 80 meters (262 feet) and occasionally venturing into deep dives reaching as low as 616 meters (2,021 feet). These deeper descents are believed to serve various purposes, including navigational guidance, communication with fellow humpback whales, and facilitation of feeding activities.[22] Dives typically do not exceed five minutes during the summer but are normally 15–20 minutes during the winter.[13] As it dives, a humpback typically raises its tail fluke, exposing the underside.[10]
الغذاء
Humpback whales feed from spring to fall. They are generalist feeders, their main food items being krill, copepods, other plankton and small schooling fish. The most common krill species eaten in the southern hemisphere is the Antarctic krill. Further north, the northern krill and various species of Euphausia and Thysanoessa are consumed. Fish prey include herring, capelin, sand lances and Atlantic mackerel.[10][13] Like other rorquals, humpbacks are "gulp feeders", swallowing prey in bulk, while right whales and bowhead whales are skimmers.[17] The whale increases its mouth gape by expanding the grooves.[10] Water is pushed out through the baleen.[23] In the southern hemisphere, humpbacks have been recorded foraging in large compact gatherings numbering up to 200 individuals.[24]
Humpbacks hunt their prey with bubble-nets. A group swims in a shrinking circle while blowing air from their blowholes, capturing the prey above them in a cylinder of bubbles. They may dive up to 20 m (66 ft) performing this technique. Bubble-netting comes in two main forms; upward spirals and double loops. Upward spirals involve the whales blowing air from their blowhole continuously as they circle towards the surface, creating a spiral of bubbles. Double loops consist of a deep, long loop of bubbles that herds the prey, followed by slapping the surface and then a smaller loop that prepares the final capture. Combinations of spiraling and looping have been recorded. After the humpbacks create the "nets", the whales swim into them with their mouths gaping and ready to swallow.[23]
Using network-based diffusion analysis, one study argued that whales learned lobtailing from other whales in the group over 27 years in response to a change in primary prey.[25][26] The tubercles on the flippers stall the angle of attack, which both maximizes lift and minimizes drag (see tubercle effect). This, along with the shape of the flippers, allows the whales to make the abrupt turns necessary during bubble-feeding.[27]
المغازلة والتكاثر
Mating and breeding take place during the winter months, which is when females reach estrus and males reach peak testosterone and sperm levels.[10] Humpback whales are promiscuous, with both sexes having multiple partners.[10][28] Males will frequently trail both lone females and cow–calf pairs. These are known as "escorts", and the male that is closest to the female is known as the "principal escort", who fights off the other suitors known as "challengers". Other males, called "secondary escorts", trail further behind and are not directly involved in the conflict.[29] Agonistic behavior between males consists of tail slashing, ramming, and head-butting.[10] Males have also been observed engaging in copulation with each other.[30]
Gestation in the species lasts 11.5 months, and females reproduce every two years.[10] Humpback whale births have been rarely observed. One birth witnessed off Madagascar occurred within four minutes.[31] Mothers typically give birth in mid-winter, usually to a single calf. Calves suckle for up to a year but can eat adult food in six months. Humpbacks are sexually mature at 5–10 years, depending on the population.[10] Humpback whales possibly live over 50 years.[11]
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الأصوات
Male humpback whales produce complex songs during the winter breeding season. These vocals range in frequency between 100 Hz and 4 kHz, with harmonics reaching up to 24 kHz or more, and can travel at least 10 km (6.2 mi). Males may sing for between 4 and 33 minutes, depending on the region. In Hawaii, humpback whales have been recorded vocalizing for as long as 7 hours.[32] Songs are divided into layers; "subunits", "units", "subphrases", "phrases" and "themes". A subunit refers to the discontinuities or inflections of a sound while full units are individual sounds, similar to musical notes. A succession of units creates a subphrase, and a collection of subphrases make up a phrase. Similar-sounding phrases are repeated in a series grouped into themes, and multiple themes create a song.[33]
The function of these songs has been debated, but they may have multiple purposes. There is little evidence to suggest that songs establish dominance among males. However, there have been observations of non-singing males disrupting singers, possibly in aggression. Those who join singers are males who were not previously singing. Females do not appear to approach singers that are alone, but may be drawn to gatherings of singing males, much like a lek mating system. Another possibility is that songs bring in foreign whales to populate the breeding grounds.[32] It has also been suggested that humpback whale songs have echolocating properties and may serve to locate other whales.[34] A 2023 study found that as humpback whales numbers have recovered from whaling, singing has become less common.[35]
Whale songs are similar among males in a specific area. Males may alter their songs over time, and others in contact with them copy these changes.[33] They have been shown in some cases to spread "horizontally" between neighboring populations throughout successive breeding seasons.[36] In the northern hemisphere, songs change more gradually while southern hemisphere songs go through cyclical "revolutions".[37]
Humpback whales are reported to make other vocalizations. "Snorts" are quick low-frequency sounds commonly heard among animals in groups consisting of a mother–calf pair and one or more male escort groups. These likely function in mediating interactions within these groups. "Grumbles" are also low in frequency but last longer and are more often made by groups with one or more adult males. They appear to signal body size and may serve to establish social status. "Thwops" and "wops" are frequency modulated vocals, and may serve as contact calls both within and between groups. High-pitched "cries" and "violins" and modulated "shrieks" are normally heard in groups with two or more males and are associated with competition. Humpback whales produce short, low-frequency "grunts" and short, modulated "barks" when joining new groups.[38]
الافتراس
Visible scars indicate that orcas prey upon juvenile humpbacks.[17] A 2014 study in Western Australia observed that when available in large numbers, young humpbacks can be attacked and sometimes killed by orcas. Moreover, mothers and (possibly related) adults escort calves to deter such predation. The suggestion is that when humpbacks suffered near-extinction during the whaling era, orcas turned to other prey but are now resuming their former practice.[39] There is also evidence that humpback whales will defend against or mob killer whales who are attacking either humpback calves or juveniles as well as members of other species, including seals. The humpback's protection of other species may be unintentional, a "spillover" of mobbing behavior intended to protect members of its species. The powerful flippers of humpback whales, often infested with large, sharp barnacles, are formidable weapons against orcas. When threatened, they will thresh their flippers and tails keeping the orcas at bay.[40]
The great white shark is another confirmed predator of the humpback whale. In 2020, Marine biologists Dines and Gennari et al., published a documented incident of a group of great white sharks exhibiting pack-like behavior to attack and kill a live adult humpback whale.[41] A second incident regarding great white sharks killing humpback whales was documented off the coast of South Africa. The shark recorded instigating the attack was a female nicknamed "Helen". Working alone, the shark attacked a 33 ft (10 m) emaciated and entangled humpback whale by attacking the whale's tail to cripple and bleed the whale before she managed to drown the whale by biting onto its head and pulling it underwater.[42][43]
النطاق
Humpback whales are found worldwide, except for some areas at the equator and High Arctic and some enclosed seas.[11] The furthest north they have been recorded is at 81°N around northern Franz Josef Land.[44] They are usually coastal and tend to congregate in waters within continental shelves. Their winter breeding grounds are located around the equator; their summer feeding areas are found in colder waters, including near the polar ice caps. Humpbacks go on vast migrations between their feeding and breeding areas, often crossing the open ocean. The species has been recorded traveling up to 8,000 km (5,000 mi) in one direction.[11] An isolated, non-migratory population feeds and breeds in the northern Indian Ocean, mainly in the Arabian Sea around Oman.[45] This population has also been recorded in the Gulf of Aden, the Persian Gulf, and off the coasts of Pakistan and India.[46]
In the North Atlantic, there are two separate wintering populations, one in the West Indies, from Cuba to northern Venezuela, and the other in the Cape Verde Islands and northwest Africa. During summer, West Indies humpbacks congregate off New England, eastern Canada, and western Greenland, while the Cape Verde population gathers around Iceland and Norway. There is some overlap in the summer ranges of these populations, and West Indies humpbacks have been documented feeding further east.[45] Whale visits into the Gulf of Mexico have been infrequent but have occurred in the gulf historically.[47] They were considered to be uncommon in the Mediterranean Sea, but increased sightings, including re-sightings, indicate that more whales may colonize or recolonize it in the future.[48]
The North Pacific has at least four breeding populations: off Mexico (including Baja California and the Revillagigedos Islands), Central America, the Hawaiian Islands, and both Okinawa and the Philippines. The Mexican population forages from the Aleutian Islands to California, particularly the Bering Sea, northern and western Gulf of Alaska, southern British Columbia to northern Washington State, and Oregon to California. During the summer, Central American humpbacks are found only off Oregon and California. In contrast, Hawaiian humpbacks have a wide feeding range but most travel to southeast Alaska and northern British Columbia. The wintering grounds of the Okinawa/Philippines population are mainly around the Russian Far East. There is some evidence for a fifth population somewhere in the northwestern Pacific. These whales are recorded to feed off the Aleutians with a breeding area somewhere south of the Bonin Islands.[45]
نصف الكرة الجنوبي
في نصف الكرة الجنوبي، humpback whales are divided into seven breeding stocks, some of which are further divided into sub-structures. These include the southeastern Pacific (stock G), southwestern Atlantic (stock A), southeastern Atlantic (stock B), southwestern Indian Ocean (stock C), southeastern Indian Ocean (stock D), southwestern Pacific (stock E), and the Oceania stock (stocks E–F).[45] Stock G breeds in tropical and subtropical waters off the west coast of Central and South America and forages along the west coast of the Antarctic Peninsula, the South Orkney Islands and to a lesser extent the Tierra del Fuego of southern Chile. Stock A winters off Brazil and migrates to summer grounds around South Georgia and the South Sandwich Islands. Some stock A individuals have also been recorded off the western Antarctic Peninsula, suggesting an increased blurring of the boundaries between the feeding areas of stocks A and G.[49]
Stock B breeds on the west coast of Africa and is further divided into Bl and B2 subpopulations, the former ranging from the Gulf of Guinea to Angola and the latter ranging from Angola to western South Africa. Stock B whales have been recorded foraging in waters to the southwest of the continent, mainly around Bouvet Island.[50] Comparison of songs between those at Cape Lopez and Abrolhos Archipelago indicate that trans-Atlantic mixings between stock A and stock B whales occur.[51] Stock C whales winter around southeastern Africa and surrounding waters. This stock is further divided into C1, C2, C3, and C4 subpopulations; C1 occurs around Mozambique and eastern South Africa, C2 around the Comoro Islands, C3 off the southern and eastern coast of Madagascar and C4 around the Mascarene Islands. The feeding range of this population is likely between coordinates 5°W and 60°E and under 50°S.[45][50] There may be overlap in the feeding areas of stocks B and C.[50]
Stock D whales breed off the western coast of Australia, and forage in the southern region of the Kerguelen Plateau.[52] Stock E is divided into E1, E2, and E3 stocks.[45] E1 whales have a breeding range off eastern Australia and Tasmania; their main feeding range is close to Antarctica, mainly within 130°E and 170°W.[53] The Oceania stock is divided into the New Caledonia (E2), Tonga (E3), Cook Islands (F1) and French Polynesia (F2) subpopulations. This stock's feeding grounds mainly range from around the Ross Sea to the Antarctic Peninsula.[54]
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العلاقة بالبشر
صيد الحيتان
Humpback whales were hunted as early as the late 16th century.[2] They were often the first species to be harvested in an area due to this coastal distribution.[10] North Pacific kills alone are estimated at 28,000 during the 20th century.[12] In the same period, over 200,000 humpbacks were taken in the Southern Hemisphere.[10] North Atlantic populations dropped to as low as 700 individuals.[12] In 1946, the International Whaling Commission (IWC) was founded to oversee the industry. They imposed hunting regulations and created hunting seasons. To prevent extinction, IWC banned commercial humpback whaling in 1966. By then, the global population had been reduced to around 5,000.[55] The Soviet Union deliberately under-recorded its catches; the Soviets reported catching 2,820 between 1947 and 1972, but the true number was over 48,000.[56]
As of 2004, hunting was restricted to a few animals each year off the Caribbean island of Bequia in Saint Vincent and the Grenadines.[57] The take is not believed to threaten the local population. Japan had planned to kill 50 humpbacks in the 2007/08 season under its JARPA II research program. The announcement sparked global protests.[58] After a visit to Tokyo by the IWC chair asking the Japanese for their co-operation in sorting out the differences between pro- and anti-whaling nations on the commission, the Japanese whaling fleet agreed to take no humpback whales during the two years it would take to reach a formal agreement.[59] In 2010, the IWC authorized Greenland's native population to hunt a few humpback whales for the following three years.[60]
حالة الحفاظ
As of 2018, the IUCN Red List lists the humpback whale as least-concern, with a worldwide population of around 135,000 whales, of which around 84,000 are mature individuals, and an increasing population trend.[2] Prior to 2008, the IUCN listed the species as vulnerable.[61] Regional estimates are around 13,000 in the North Atlantic, 21,000 in the North Pacific, and 80,000 in the southern hemisphere. For the isolated population in the Arabian sea, only around 80 individuals remain,[62] and this population is considered to be endangered. In most areas, humpback whale populations have recovered from historic whaling, particularly in the North Pacific.[11] Such recoveries have led to the downlisting of the species' threatened status in the United States, Canada, and Australia.[61][63] In Costa Rica, Ballena Marine National Park was established for humpback protection.[64]
Humpbacks still face various other human-made threats, including entanglement by fishing gear, vessel collisions, human-caused noise and traffic disturbance, coastal habitat destruction, and climate change.[11] Like other cetaceans, humpbacks can be injured by excessive noise. In the 19th century, two humpback whales were found dead near repeated oceanic sub-bottom blasting sites, with traumatic injuries and fractures in the ears.[65] Saxitoxin, a paralytic shellfish poisoning from contaminated mackerel, has been implicated in humpback whale deaths.[66] While oil ingestion is a risk for whales, a 2019 study found that oil did not foul baleen and instead was easily rinsed by flowing water.[67]
Whale researchers along the Atlantic Coast report that there have been more stranded whales with signs of vessel strikes and fishing gear entanglement in recent years than ever before. The NOAA recorded 88 stranded humpback whales between January 2016 and February 2019. This is more than double the number of whales stranded between 2013 and 2016. Because of the increase in stranded whales, NOAA declared an unusual mortality event in April 2017. Virginia Beach Aquarium's stranding response coordinator, Alexander Costidis, stated the conclusion that the two causes of these unusual mortality events were vessel interactions and entanglements.[68]
مشاهدة الحيتان
Much of the growth of commercial whale watching was built on the humpback whale. The species' highly active surface behaviors and tendency to become accustomed to boats have made them easy to observe, particularly for photographers. In 1975, humpback whale tours were established in New England and Hawaii.[69] This business brings in a revenue of $20 million per year for Hawaii's economy.[70] While Hawaiian tours have tended to be commercial, New England and California whale watching tours have introduced educational components.[69]
There are many commercial whale-watching operations on both the humpback's summer and winter ranges:
شمال الأطلسي | North Pacific | Southern Hemisphere | |
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Summer | New England, Nova Scotia and Newfoundland, the northern St. Lawrence River, the Snaefellsnes peninsula in the west of Iceland | كاليفورنيا, ألاسكا, Oregon, Washington, British Columbia | أنتارتيكا, Bahía Solano and Nuquí in Colombia |
Winter | Samaná Province of the Dominican Republic, the Bay of Biscay France, | هاوائي, Baja, the Bahía de Banderas off Puerto Vallarta | Sydney, Byron Bay north of Sydney, Hervey Bay north of Brisbane, North and East of Cape Town, New Zealand, the Tongan islands, |
As with other cetacean species, however, a mother whale is generally extremely protective of her infant, and places herself between any boat and her calf before moving quickly away from the vessel. Skilled tour operators avoid stressing the mother.
الإعلام
هل لديك مشكلة في تشغيل هذه الملفات؟ انظر مساعدة الوسائط.
ميگالو
An albino humpback whale that travels up and down the east coast of Australia became famous in local media because of its rare, all-white appearance. Migaloo is the only known Australian all-white specimen,[71] and is a true albino.[72] First sighted in 1991, the whale was named for an indigenous Australian word for "white fella". To prevent sightseers from approaching dangerously close, the Queensland government decreed a 500-m (1600-ft) exclusion zone around him.[73]
Migaloo was last seen in June 2014 along the coast of Cape Byron in Australia. Migaloo has several physical characteristics that can be identified; his dorsal fin is somewhat hooked, and his tail flukes have a unique shape, with edges that are spiked along the lower trailing side.[74] In July 2022, concerns arose after a white whale washed up on the shores of Mallacoota beach, however after genetic testing, and noting that the carcass was of a female whale while Migaloo is male, it was confirmed by experts to not be Migaloo.[75][76]
همفرى
In 1985, Humphrey swam into San Francisco Bay and then up the Sacramento River towards Rio Vista.[77] Five years later, Humphrey returned and became stuck on a mudflat in San Francisco Bay immediately north of Sierra Point below the view of onlookers from the upper floors of the Dakin Building. He was twice rescued by the Marine Mammal Center and other concerned groups in California.[78] He was pulled off the mudflat with a large cargo net and the help of the US Coast Guard. Both times, he was successfully guided back to the Pacific Ocean using a "sound net" in which people in a flotilla of boats made unpleasant noises behind the whale by banging on steel pipes, a Japanese fishing technique known as oikami. At the same time, the attractive sounds of humpback whales preparing to feed were broadcast from a boat headed towards the open ocean.[79]
دلتا و دون
هوامش
- ^ قالب:MSW3 Cetacea
- ^ أ ب ت خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةiucn
- ^ أ ب خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةWJ
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةLiddellScott2015
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةArnason_etal_2018
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةReeves
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةHatch
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةsubspecies
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةPomilla
- ^ أ ب ت ث ج ح خ د ذ ر ز س ش ص ض ط خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةPerrin
- ^ أ ب ت ث ج ح خ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةJefferson
- ^ أ ب ت ث خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةplan
- ^ أ ب ت ث ج ح خ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةmammalian
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةtubercles
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةKatonaWhitehead
- ^ خطأ استشهاد: وسم
<ref>
غير صحيح؛ لا نص تم توفيره للمراجع المسماةKaufmanSmultea
- ^ أ ب ت ث خطأ استشهاد: وسم
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- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةAbrolhos
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةMobley
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةDeakos
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةIwata
- ^ Derville, Solène; Torres, Leigh G.; Zerbini, Alexandre N.; Oremus, Marc; Garrigue, Claire (2020-03-17). "Horizontal and vertical movements of humpback whales inform the use of critical pelagic habitats in the western South Pacific". Scientific Reports (in الإنجليزية). 10 (1): 4871. Bibcode:2020NatSR..10.4871D. doi:10.1038/s41598-020-61771-z. ISSN 2045-2322. PMC 7078318. PMID 32184421.
- ^ أ ب خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةwhaleculture
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةFish
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةClapham97
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةHerman
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةStack
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةshortnote
- ^ أ ب خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةHerman2017
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةCholewiak2012
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةsonic
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةDunlop
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةpmid21497089
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةZanberg
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةRESGA
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةmobbing
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةDinesandGennari
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةIndependent
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةexpress
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةFranz
- ^ أ ب ت ث ج ح خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةNOAA
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةRosenbaum
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةsouthwestPacific
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةbaker
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةRecplan91
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةBBC1
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةPress
- ^ أ ب خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةMarinepolicy
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةPerrin2015
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةabc
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةUTP
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةketten
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةDieraufGulland2001
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةroyal
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةnatgeo
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةhawaii
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةmigaloo
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةjhered
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةSUP
- ^ Thomas, Kerrin; Hook, Mim (16 July 2022). "White whale washes up on Mallacoota beach". ABC News. Australian Broadcasting Corporation. Retrieved 16 July 2022.
- ^ "Experts confirm white whale carcass on Victorian beach is not Migaloo". Australian Associated Press. The Guardian. 17 July 2022. Retrieved 17 July 2022.
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةkay
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةtokuda
- ^ خطأ استشهاد: وسم
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غير صحيح؛ لا نص تم توفيره للمراجع المسماةKnapp1993
المصادر
كتب
- Clapham, Phil. (1996). Humpback Whales. ISBN 0-948661-87-9
- Clapham, Phil (2002). "Humpback Whale". Encyclopedia of Marine Mammals. San Diego: Academic Press. ISBN 0-12-551340-2.
- Dawbin, W. H. The seasonal migratory cycle of humpback whales. In K.S. Norris (ed), Whales, Dolphins and Porpoises. University of California Press.
- Dawes, John (2008). Exploring the World of Aquatic Life. Infobase Publishing. ISBN 978-1-60413-255-7. Retrieved 2011-01-08.
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suggested) (help) - Evans, Peter G. H., and Juan Antonio. Raga. Marine Mammals: Biology and Conservation. New York: Kluwer Academic/Plenum, 2001. Print.
- Reeves, Stewart, Clapham and Powell. (2002) National Audubon Society Guide to Marine Mammals of the World. Knopf. ISBN 0-375-41141-0
مقالات صحفية
- Best, P. B. (1993). "Increase rates in severely depleted stocks of baleen whales". ICES Journal of Marine Science. 50: 169–186. doi:10.1006/jmsc.1993.1018.
- Smith, T.D.; Allen, J.; Clapham, P.J.; Hammond, P.S.; Katona, S.; Larsen, F.; Lien, J.; Mattila, D.; Palsboll, P.J.; et al. (1999). "An ocean-basin-wide mark-recapture study of the North Atlantic humpback whale". Marine Mammal Science. 15: 1–32. doi:10.1111/j.1748-7692.1999.tb00779.x.
{{cite journal}}
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(help)
وصلات خارجية
- General
- US National Marine Fisheries Service Humpback Whale web page
- ARKive - images and movies of the humpback whale (Megaptera novaeangliae).
- Photos and information about humpback whales and other cetaceans in Mayumba National Park
- Humpbacks of Hervey Bay, Queensland, Australia
- The Dolphin Institute Whale Resource Guide and scientific publications
- Humpback Whale Gallery (Silverbanks)
- (بالفرنسية)Humpback whale videos
- Book about the two humpback whales lost in the Sacramento River in May 2007 -
- The Humpback Whales of Hervey Bay
- Epic humpback whale battle filmed
- أغاني الحوت الأحدب
- The Whalesong Project
- Article from PHYSORG.com on the complex syntax of whalesong phrases
- Analysis of the form and function of the humpback whale song from the University at Buffalo
- الحفاظ
- Short description is different from Wikidata
- أنواع القائمة الحمراء غير المهددة
- Articles containing Ancient Greek (to 1453)-language text
- Articles with hatnote templates targeting a nonexistent page
- Articles with hAudio microformats
- CS1 errors: unsupported parameter
- مقالات ناطقة
- وصف الحيوانات في 1781
- حيتان البالين
- حيتان في استراليا
- حيوانات في جرينلاند
- حيوانات في المحيط الأطلسي
- حيوانات في المحيط الهادئ
- ثدييات بحرية في هاواي
- حيوانات ضخمة
- أجناس ثدييات أحادية النوع